First direct evidence of lion hunting and the early use of a lion pelt by Neanderthals.

During the Upper Paleolithic, lions become an important theme in Paleolithic art and are more frequent in anthropogenic faunal assemblages. However, the relationship between hominins and lions in earlier periods is poorly known and primarily interpreted as interspecies competition. Here we present new evidence for Neanderthal-cave lion interactions during the Middle Paleolithic. We report new evidence of hunting lesions on the 48,000 old cave lion skeleton found at Siegsdorf (Germany) that attest to the earliest direct instance of a large predator kill in human history. A comparative analysis of a partial puncture to a rib suggests that the fatal stab was delivered with a wooden thrusting spear. We also present the discovery of distal lion phalanges at least 190,000 old from Einhornhöhle (Germany), representing the earliest example of the use of cave lion skin by Neanderthals in Central Europe. Our study provides novel evidence on a new dimension of Neanderthal behavioral complexity.

The basal Lutetian Transgression on the Tethyan shelf of the European craton (Adelholzen beds, Eastern Alps, Germany).

The middle Eocene Adelholzen beds were deposited on the northern part of the Tethyan shelf of the European Plate. In the Eastern Alps, the Wimmern section near Teisendorf (Germany) is the only known outcrop exposing the basal unconformity of this sequence. The outcrop comprises an 8 m thick condensed transgressive shallow marine succession characterized by glauconite-rich weakly consolidated greensand and mudstone containing abundant larger benthic foraminifera. It spans the lower part of calcareous nannoplankton Zone NP15 (Sub-Zones NP15a and lower part of NP15b), planktonic foraminifera Sub-Zone E7b and Zone E8 partim and the upper part of shallow benthic Zone SBZ13. The landward migration of the paleo-shoreline was not an effect of flexural downbending of the European Plate but can be correlated to the major unconformity at the base of eustatic supercycle TA3. The onset of this sea-level rise was in the latest part of Biochron NP14b and almost coincided with the NP14b/NP15a-boundary.

Middle to Late Eocene paleoenvironmental changes in a marine transgressive sequence from the northern Tethyan margin (Adelholzen, Germany).

The northern Tethyan margin is a key region for determining environmental changes associated with the collision of continental and oceanic tectonic plates and Alpine orogeny. Herein we investigated Middle to Late Eocene neritic to bathyal sediments deposited during an interval of unstable climatic conditions. In order to quantify paleoenvironmental changes, we developed a detailed age model based on biozonations of planktic foraminifera, calcareous nannoplankton, and larger benthic foraminifera. The section at Adelholzen covers the almost complete Lutetian Stage (calcareous nannoplankton zones NP15a-16, planktic foraminifera zones E8-11, shallow benthic (foraminifera) zones SBZ13-15) and large parts of the Priabonian Stage (NP18-20, E14/15), while the intermediate Bartonian Stage (NP17) is completely missing. Foraminiferal, calcareous nannoplankton, and macrofossil assemblages were analyzed for changes in paleo-water depth, mixing and stratification, paleo-primary productivity (pPP), food supply, and bottom water oxygenation. Paleo-water depth estimates range from 50 m (middle neritic, early Lutetian) to nearly 500 m (upper bathyal, late Priabonian). The combination of assemblage composition, planktic and benthic foraminiferal accumulation rates, and derived parameters (carbon-flux to sea floor, pPP) enabled us to identify a series of distinct paleoceanographic events of at least regional significance. Such events are characterized by considerable changes in primary productivity or reduced bottom water ventilation. Calculated pPP-values indicate oligotrophic conditions throughout.